An arthropod is an invertebrate animal having an exoskeleton (external skeleton), a segmented body, and jointed appendages. Arthropods are members of the phylum Arthropoda (from Greek ?????? árthron, "joint", and ????? podós "foot", which together mean "jointed feet"), and include the insects, arachnids, crustaceans, and others. Arthropods are characterized by their jointed limbs and cuticles, which are mainly made of ?-chitin; the cuticles of crustaceans are also biomineralized with calcium carbonate. The rigid cuticle inhibits growth, so arthropods replace it periodically by molting. The arthropod body plan consists of repeated segments, each with a pair of appendages. It is so versatile that they have been compared to Swiss Army knives, and it has enabled them to become the most species-rich members of all ecological guilds in most environments. They have over a million described species, making up more than 80% of all described living animal species, and are one of only two animal groups that are very successful in dry environments – the other being the amniotes. They range in size from microscopic plankton up to forms a few meters long. Ernst Heinrich Philipp August Haeckel (16 February 1834 – 9 August 1919) was a German biologist, naturalist, philosopher, physician, professor, marine biologist, and artist who discovered, described and named thousands of new species, mapped a genealogical tree relating all life forms, and coined many terms in biology.